Ancestry.com LLC is a privately held online company based in Lehi, Utah. The largest for-profit genealogy company in the world, it operates a network of genealogical, historical record and genetic genealogy websites.

As of November 2018, the company claimed to provide access to approximately 10 billion historical records, to have 3 million paying subscribers and to have sold 14 million DNA kits to customers.

In 1990, Paul B. Allen^[4] and Dan Taggart, two Brigham Young University graduates, founded Infobases and began offering Latter-day Saints (LDS) publications on floppy disks. In 1988, Allen had worked at Folio Corporation, founded by his brother Curt and his brother-in-law Brad Pelo.

Infobases' first products were floppy disks and compact disks sold from the back seat of the founders' car.

In 1994, Infobases was named among Inc. magazine's 500 fastest-growing companies.^[5] Their first offering on CD was the LDS Collectors Edition, released in April 1995, selling for $299.95,^[6] which was offered in an online version in August 1995.^[7] Ancestry officially went online with the launch of Ancestry.com in 1996.^[8]

On January 1, 1997, Infobases' parent company, Western Standard Publishing, purchased Ancestry, Inc.,^[9] publisher of Ancestry magazine and genealogy books. Western Standard Publishing's CEO was Joe Cannon, one of the principal owners of Geneva Steel.^[10]

In July 1997, Allen and Taggart purchased Western Standard's interest in Ancestry, Inc. At the time, Brad Pelo was president and CEO of Infobases, and president of Western Standard. Less than six months earlier, he had been president of Folio Corporation, whose digital technology Infobases was using. In March 1997, Folio was sold to Open Market for $45 million.^[11] The first public evidence of the change in ownership of Ancestry magazine came with the July/August 1997 issue, which showed a newly reorganized Ancestry, Inc., as its publisher. That issue's masthead also included the first use of the Ancestry.com web address.

More growth for Infobases occurred in July 1997, when Ancestry, Inc. purchased Bookcraft, Inc., a publisher of books written by leaders and officers of the LDS Church.^[12][13] Infobases had published many of Bookcraft's books as part of its LDS Collector's Library. Pelo also announced that Ancestry's product line would be greatly expanded in both CDs and online. Alan Ashton, a longtime investor in Infobases and founder of WordPerfect, was its chairman of the board.

Allen and Taggart began running Ancestry, Inc. independently from Infobases in July 1997, and began creating one of the largest online subscription-based genealogy database services.^[14]

In April 1999, to better focus on its Ancestry.com and MyFamily.com Internet businesses, Infobases sold the Bookcraft brand name and its catalog of print books to its major competitor in the LDS book market, Deseret Book. Included in the sale were the rights to Infobases' LDS Collectors Library on CD. A year earlier, Deseret Book had released a competing product called GospeLink, and the two products were combined as a single product by Deseret Book.^[15][16]

The MyFamily.com website launched in December 1998, with additional free sites beginning in March 1999.^[17] The site generated one million registered users within its first 140 days.^[14] The company raised more than US$90 million in venture capital from investors^[14] and changed its name on November 17, 1999, from Ancestry.com, Inc. to MyFamily.com, Inc. Its three Internet genealogy sites were then called Ancestry.com, FamilyHistory.com, and MyFamily.com.^[18] Sales were about US$62 million for 2002 and US$99 million for 2003.^[19]

In March 2004, the company, which had outgrown its call center in Orem, Utah, opened a new call center, which accommodates about 700 agents at a time, in Provo.^[20] Heritage Makers was acquired by MyFamily.com in September 2005.^[21]

While the company had been offering free access to Ancestry.com at LDS Family History Centers, that service was terminated on March 17, 2007, because the company and the LDS Church were unable to reach a mutually agreeable licensing agreement. In 2010, Ancestry restored access to its site at Family History Centers.

In 2010, Ancestry sold its book publishing assets to Turner Publishing Company.^[22]

Ancestry.com became a publicly traded company on NASDAQ (symbol: ACOM) on November 5, 2009, with an initial public offering of 7.4 million shares priced at $13.50 per share, underwritten by Morgan Stanley, Bank of America, Merrill Lynch, Jefferies & Company, Piper Jaffray, and BMO Capital Markets.^[23]

In 2010, Ancestry.com expanded its domestic operations with the opening of an office in San Francisco, California, staffed with brand new engineering, product, and marketing teams geared toward developing some of Ancestry's cutting-edge technology and services. In 2011, Ancestry launched an Android and iOS app.^[24][25]

In December 2011, Ancestry.com moved the Social Security Death Index search behind a paywall and stopped displaying the Social Security information of people who had died within the past 10 years, because of identity theft concerns.^[26]

In March 2012, Ancestry.com acquired the collection of DNA assets from GeneTree.^[27]

In September 2012, Ancestry.com expanded its international operations with the opening of its European headquarters in Dublin, Ireland. The Dublin office includes a new call centre for international customers, as well as product, marketing, and engineering teams.^[28][29]

In October 2012, Ancestry.com agreed to be acquired by a private equity group consisting of Permira Advisers LLP, members of Ancestry.com's management team, including CEO Tim Sullivan and CFO Howard Hochhauser, and Spectrum Equity, for $32 per share or around $1.6 billion.^[30] At the same time, Ancestry.com purchased a photo digitization and sharing service called 1000Memories.^[31]

On July 16, 2015, Ancestry launched AncestryHealth, and announced the appointment of Cathy A. Petti as its Chief Health Officer.^[32]

In April 2016 GIC Private Limited (a sovereign wealth fund owned by the Government of Singapore) and Silver Lake (a private equity fund manager) bought equity stakes in Ancestry.com.^[33] The estimated market value of Ancestry.com in 2017 was more than $3 billion.^[34]

In November 2018 Ancestry claimed to have over 10 billion digitised records and over three million paying customers.^[35]

AncestryDNA is a subsidiary of Ancestry LLC.

AncestryDNA offers a direct-to-consumer genealogical DNA test.^[36] Consumers provide a sample of their DNA to the company for analysis. AncestryDNA then uses DNA sequences to infer family relationships with other Ancestry DNA users and to provide what it calls an "ethnicity estimate". Previously, Ancestry.com also offered paternal Y-chromosome DNA and maternal mitochondrial DNA tests, but those were discontinued in June 2014.^[37] The company describes the technical process of testing in a scientific white paper. In May 2019 the company claimed that their database contained 15 million completed DNA kits bought by customers.^[38][35][39]

Ancestry DNA is commonly used for donor conceived persons to find their biological siblings and in some cases their sperm or egg donor.^[40]

The testing itself is performed by Quest Diagnostics.^[41]

On September 30, 2013, Ancestry.com announced its acquisition of Find a Grave. Site editor Jim Tipton said of the purchase that Ancestry.com had "been linking and driving traffic to the site for several years. Burial information is a wonderful source for people researching their family history". Ancestry.com launched a mobile app in March 2014.^[42]

In 2012, Ancestry spun off its digitized online newspaper components into a standalone service Newspapers.com with newspaper.com only pricing as well as a bundled Ancestry.com pricing.

Before the Newspapers.com launch, Ancestry.com acquired the following newspaper-oriented components, including scanning and digital technologies and posting on the web:

The website's principal competitor is newspaperarchive.com which claims it has online newspapers dating from 1607 worldwide and its index in June 2018 includes 9,829 newspapers.^[49] Both websites have similar models for increasing their databases: striking deals with libraries, publishers and historical organizations to scan the publications for free to include in their database. Participants note that the process of free scanning is easier, cheaper and quicker to get their publications online rather than working through the government operated National Digital Newspaper Program.^[50]

RootsWeb, acquired by Ancestry in June 2000, is a free genealogy community that uses online forums, mailing lists, and other resources to help people research their family history.

Users can upload GEDCOM files of their information for others to search at the WorldConnect portion of the site. Trees uploaded to WorldConnect are searchable at both the RootsWeb and Ancestry websites.

On December 20, 2017, a file containing 300,000 RootsWeb user names, passwords, and email addresses was exposed to the internet.

The 300,000 records were from RootsWeb surname list service with 55,000 of those records were also Ancestry.com login credentials.^[51]

In 2013, Ancestry partnered with FamilySearch. The two genealogical organisations agreed to co-operate in digitizing records and to allow members of the Church of Jesus Christ of Latter Day Saints to have free Ancestry accounts. ^[64]

In 2015, Ancestry partnered with the Google subsidiary, Calico, to focus on longevity research and therapeutics, in an effort to investigate human heredity of lifespan.^[65][66]

After the arrest of the accused Golden State Killer, Ancestry.com and 23andMe made a data policy stating that they won't allow their DNA profiles to be used for crime solving unless pressured to do so by law enforcement, as they believe it violates users' privacy. This is despite the fact that Ancestry.com's database was used in 2017 to solve the murder of Jane Britton.^[67][68]

In 2019, Ancestry.com was a awarded a German Big Brother Award, a negative award, "for exploiting an interest in genealogy to entice people into submitting saliva samples … to pile up a treasure trove of genome data for commercial research, because that is their actual business model."^[69]